Some Straw-Hat Notes;
Came across some goodies looking at the link Shed sent earlier about tree injections. One door opens another room full of doors.
WIKI;
Xylem is one of the two types of transport
tissue in
vascular plants,
phloem being the other. The basic function of xylem is to transport
water from roots to stems and leaves, but it also transports
nutrients.
[1][2] The word "xylem" is derived from the
Greek word ξύλον (
xylon), meaning "wood"; the best-known xylem tissue is
wood, though it is found throughout a plant.
[3] The term was introduced by
Carl Nägeli in 1858.
The most distinctive xylem
cells are the long tracheary elements that transport water.
Tracheids and
vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called
vessels.
[6]
Xylem also contains two other cell types:
parenchyma and
fibers.
[7]
Xylem can be found:
in
vascular bundles, present in non-woody plants and non-woody parts of woody plants
in secondary xylem, laid down by a
meristem called the
vascular cambium in woody plants
as part of a
stelar arrangement not divided into bundles, as in many
ferns.
In transitional stages of plants with
secondary growth, the first two categories are not mutually exclusive, although usually a vascular bundle will contain
primary xylem only. The branching pattern exhibited by xylem follows
Murray's law.
The xylem, vessels and tracheids of the roots, stems and leaves are interconnected to form a continuous system of water-conducting channels reaching all parts of the plants. The system transports water and soluble mineral nutrients from the roots throughout the plant. It is also used to replace water lost during
transpiration and photosynthesis. Xylem
sap consists mainly of water and inorganic ions, although it can also contain a number of organic chemicals as well. The transport is passive, not powered by energy spent by the
tracheary elements themselves, which are dead by maturity and no longer have living contents. Transporting sap upwards becomes more difficult as the height of a plant increases and upwards transport of water by xylem is considered to limit the maximum height of trees.
[11] Three phenomena cause xylem sap to flow:
Pressure flow hypothesis: Sugars produced in the leaves and other green tissues are kept in the phloem system, creating a
solute pressure differential versus the xylem system carrying a far lower load of solutes- water and minerals. The phloem pressure can rise to several MPa,
[12] far higher than atmospheric pressure. Selective inter-connection between these systems allows this high solute concentration in the phloem to draw xylem fluid upwards by negative pressure.
Transpirational pull: Similarly, the
evaporation of
water from the surfaces of
mesophyll cells to the atmosphere also creates a negative pressure at the top of a plant. This causes millions of minute
menisci to form in the mesophyll cell wall. The resulting
surface tension causes a negative pressure or
tension in the xylem that pulls the water from the roots and soil.
Root pressure: If the
water potential of the root cells is more negative than that of the
soil, usually due to high concentrations of
solute, water can move by
osmosis into the root from the soil. This causes a positive pressure that forces sap up the xylem towards the leaves. In some circumstances, the sap will be forced from the leaf through a
hydathode in a phenomenon known as
guttation. Root pressure is highest in the morning before the stomata open and allow transpiration to begin. Different plant species can have different root pressures even in a similar environment; examples include up to 145 kPa in
Vitis riparia but around zero in
Celastrus orbiculatus.
[13]
The primary force that creates the
capillary action movement of water upwards in plants is the adhesion between the water and the surface of the xylem conduits.
[14][15] Capillary action provides the force that establishes an equilibrium configuration, balancing gravity. When transpiration removes water at the top, the flow is needed to return to the equilibrium.
Transpirational pull results from the evaporation of water from the surfaces of
cells in the
leaves. This evaporation causes the surface of the water to recess into the
pores of the
cell wall. By
capillary action, the water forms concave
menisci inside the pores. The high surface tension of water pulls the concavity outwards, generating enough
force to lift water as high as a hundred meters from ground level to a
tree's highest branches.
Transpirational pull requires that the vessels transporting the water be very small in diameter; otherwise,
cavitation would break the water column. And as water
evaporates from leaves, more is drawn up through the plant to replace it. When the water pressure within the xylem reaches extreme levels due to low water input from the roots (if, for example, the soil is dry), then the gases come out of solution and form a bubble – an
embolism forms, which will spread quickly to other adjacent cells, unless
bordered pits are present (these have a plug-like structure called a torus, that seals off the opening between adjacent cells and stops the embolism from spreading).
Cohesion-tension theory
The
cohesion-tension theory is a
theory of
intermolecular attraction that explains the process of
water flow upwards (against the force of
gravity) through the xylem of
plants. It was proposed in 1894 by
John Joly and
Henry Horatio Dixon.
[16][17] Despite numerous objections,
[18][19] this is the most widely accepted theory for the transport of water through a plant's vascular system based on the classical research of Dixon-Joly (1894), Eugen Askenasy (1845–1903) (1895),
[20][21] and Dixon (1914,1924).
[22][23]
Water is a
polar molecule. When two water molecules approach one another, the slightly negatively charged
oxygen atom of one forms a
hydrogen bond with a slightly positively charged
hydrogen atom in the other. This attractive force, along with other
intermolecular forces, is one of the principal factors responsible for the occurrence of
surface tension in liquid water. It also allows plants to draw water from the root through the xylem to the leaf.
Water is constantly lost through transpiration from the leaf. When one water molecule is lost another is pulled along by the processes of cohesion and tension. Transpiration pull, utilizing
capillary action and the inherent surface tension of water, is the primary mechanism of water movement in plants. However, it is not the only mechanism involved. Any use of water in leaves forces water to move into them.
Transpiration in leaves creates tension (differential pressure) in the cell walls of
mesophyll cells. Because of this tension, water is being pulled up from the roots into the leaves, helped by
cohesion (the pull between individual water molecules, due to
hydrogen bonds) and
adhesion (the stickiness between water molecules and the
hydrophilic cell walls of plants). This mechanism of water flow works because of
water potential (water flows from high to low potential), and the rules of simple
diffusion.
[24]
Over the past century, there has been a great deal of research regarding the mechanism of xylem sap transport; today, most plant scientists continue to agree that the
cohesion-tension theory best explains this process, but multiforce theories that hypothesize several alternative mechanisms have been suggested, including longitudinal cellular and xylem
osmotic pressure gradients, axial potential gradients in the vessels, and gel- and gas-bubble-supported interfacial gradients
This might take me a while to decarb all this info into stuff I need to know, and cipher ways to use the knowledge. Keeps my mind busy, even if just chasing ideas.